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The UCI School of Medicine offers graduate study leading to a Ph.D. degree in Biomedical Sciences. Five participating basic science departments train students in biomedical research and critical inquiry with the goal of a research-based career in academia, biomedical technology and related fields.   

Entry into the Ph.D. programs is primarily through the UCI Gateway Programs (CMB and INP). Direct admission is on a case-by-case basis. Students in the Medical Scientist Training Program (MSTP) may also elect to pursue their Ph.D. in Biomedical Sciences.  

About the UCI Gateway Programs:

  • Cellular & Molecular Biosciences (CMB) cmb.uci.edu
  • Interdepartmental Neuroscience Program (INP) https://inp.uci.edu

Among the first-year requirements, students supervised by a faculty advisor take coursework and identify labs for rotation with the intention of selecting a potential thesis advisor, and must pass a preliminary qualifying examination.

Application information is found on the Admission tab and at https://grad.uci.edu . 

About the Medical Scientist Training Program (MSTP):

The MSTP is an M.D./Ph.D. program that trains individuals who can understand and contribute to advances in research, and can also apply those advances to the field of medical care. Website information is found at  https://mstp.uci.edu/ . The program is administered by Joanne Ham at [email protected] and directed by Dr. Alan Goldin at [email protected] .

Departments Offering the Ph.D. in Biomedical Sciences:

  • Anatomy & Neurobiology
  • Biological Chemistry
  • Microbiology & Molecular Biology (Medical Microbiology)
  • Pathology & Laboratory Medicine (Experimental Pathology)
  • Physiology & Biophysics

Programs are administered by each department. The School of Medicine Office of Graduate Studies oversees and enhances student and program efforts. Questions may be directed to the Director of Graduate Studies, Leora Fellus at [email protected] , or Associate Dean for Graduate and Postdoctoral Studies, Dr. Peter Donovan at [email protected] .

Admission Requirements and Application for Gateway Programs and Direct Admission: 

To apply via the CMB or INP Gateway Programs, carefully review program and admission requirements and deadlines.  

  • Interdepartmental Neuroscience Program (INP):  https://inp.uci.edu/admissions-and-application/
  • Cellular & Molecular Biosciences (CMB):  https://cmb.uci.edu/admissions/

Consult each program and the UCI Graduate Division website for information and instructions:

  • https://www.grad.uci.edu/admissions/index.php
  • https://www.grad.uci.edu/admissions/applying-to-uci/index.php
  • Online Application:  https://apply.grad.uci.edu/apply/

Admission Requirements and Application for the MSTP:

Carefully review program, admission requirements and deadlines as well as qualifications and eligibility requirements for the M.D./Ph.D. program:  https://mstp.uci.edu/admissions/application-process/ .

Application for qualifying individuals is made through the American Medical College Application Service (AMCAS).

Questions about admission should be directed to program faculty and staff.

Master of Science in Biomedical Sciences

The Master of Science degree in Biomedical Sciences may be completed by submission of a research thesis (plan I) or by course-work and a comprehensive exam (plan II).

Plan I: Thesis Plan. The student is required to complete at least four didactic graduate courses (16 units) offered by the department, and elective course work with an additional 8 units of graduate or upper-division undergraduate course work. In addition, the student will typically take additional seminar courses during the graduate study. Students in the M.S. program may be employed as teaching assistants, but units earned through enrollment in University Teaching (399) may not be counted toward degree completion. The student engages in thesis research with a faculty thesis advisor, and will prepare and submit a dissertation to the thesis committee. The final examination is an oral presentation of the thesis to the committee. The normative time to degree is two years for the thesis M.S. degree.

Plan II: Comprehensive Examination Plan. The plan II M.S. degree is awarded based on completion of at least 36 units of course work and a satisfactory completion of a comprehensive exam. The student is required to complete at least 16 units (four courses) of didactic graduate course work offered by the department. In addition, the student will take up to 12 units of research. An additional 8 units or more of elective course work will be completed from other graduate courses offered by the department. A maximum of 4 units of upper-division undergraduate courses may be included in the program with the approval of the Associate Dean for Graduate Studies. Students in the M.S. program may be employed as teaching assistants, but units earned through enrollment in University Teaching (399) may not be counted toward degree completion. The comprehensive exam will be administered by a committee of at least three departmental faculty, and may include written and oral sections. The comprehensive exam format will include a research presentation and may include additional portions such as a research proposal, presentation of a project, or other components. The normative time to degree is one year for the M.S. degree by comprehensive exam.

Doctor of Philosophy

Comprehensive Examination-First Year. The student must pass comprehensive oral or written examinations at the discretion of the department. The examination is generally taken at the end of the first year of graduate study.

Advancement to Candidacy. The advancement to candidacy examination is taken before the end of the third year of graduate study. The student is expected to have identified an important and tractable dissertation research topic. A faculty committee for the advancement to candidacy examination is proposed by the faculty mentor in consultation with the student, and approved by the Department Graduate Advisor. A majority of the committee must hold primary or joint appointments in the student’s department.

Once this examination is completed, the student is advanced to candidacy for the doctoral degree and is expected to complete the degree within three years. The student must submit a dissertation on this research and defend the thesis in an oral examination during the final year of graduate study. The normative time for completion of the Ph.D. is five years, and the maximum time permitted is seven years.

Graduate-student status or consent of instructor is a prerequisite for all 200–299 courses.

  • Anatomy and Neurobiology
  • Experimental Pathology
  • Microbiology and Molecular Genetics
  • Physiology and Biophysics

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2023-2024 Catalogue

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Molecular and Cell Biology

Doctoral Program

The Department of Molecular & Cell Biology at the University of California, Berkeley offers a Ph.D. program focused on the molecular mechanisms inherent to life. This program integrates research with a modern training curricula, teaching, and career mentorship. Our Department is highly interdisciplinary - comprising the Divisions of Cell Biology, Development & Physiology, Immunology and Molecular Medicine, Molecular Therapeutics, Biochemistry, Biophysics & Structural Biology, and Genetics, Genomics, and Development – and this is reflected in our students and training. The program is also highly collaborative with related programs and Institutes on campus, thus allowing students the flexibility to explore all aspects of modern biological research. Please click on the links below to learn more about our areas of research on the main department website or use the menu at the top to navigate to areas of interest within the graduate program.

Cell Biology, Development & Physiology

Immunology and molecular medicine, neurobiology, biochemistry, biophysics & structural biology, molecular therapeutics, genetics, genomics, evolution, and development.

WHAT ARE YOU LOOKING FOR?

Key searches, phd programs, phd programs in biomedical and biological sciences (pibbs).

Portrait of five PIBBS students.

The goal of PIBBS is to recruit top caliber Ph.D. students in the biomedical and biological sciences to USC and to guide them in the identification of a mentor, lab, and research project that will provide rigorous scientific training, an outstanding Ph.D. thesis, and a strong foundation for a variety of science-related careers. The following interdisciplinary Ph.D. programs are part of PIBBS.

Cancer Biology & Genomics (CBG)

CBG capitalizes on our Norris Comprehensive Cancer Center , one of the oldest National Cancer Institute-designated Comprehensive Cancer Centers in the United States. A Comprehensive Cancer Center designation requires that clinical treatment of patients and scientific research are carried out side by side. The Norris has strength in all areas of cancer research, including clinical and basic research and epidemiology.

Development, Stem Cells, & Regenerative Medicine (DSR)

DSR benefits from our wonderful Center for Regenerative Medicine and our strength in stem cell research, tissue engineering and regeneration.

Infectious Diseases, Immunology & Pathogenesis (IDIP)

IDIP is a program focused on translating bench research to the bedside by investigating human organ systems in health and disease. It takes advantage of our outstanding clinical and basic scientists and the exciting interdisciplinary and collaborative atmosphere at USC.

Medical Biophysics (MBPH)

MBPH is a program seeks to answer important biomedical questions using all aspects of biophysics, with an emphasis on structural biology and biophysical methods and theories. Some faculty members of this program are located on the University Park (undergraduate) USC campus.

Molecular Medicine (MMED)

The Molecular Medicine PhD Program offers comprehensive research training at the intersection of medicine and molecular biology, equipping students with practical skills and a collaborative environment to make significant contributions to medical advancements.

Neuromedicine (NEUM)

The NEUM PhD program at Keck School of Medicine trains researchers to address neurological and neurodegenerative diseases through interdisciplinary approaches, equipping them with knowledge in disease biology, pathophysiology, model systems and data science, and the skills to conduct rigorous basic and translational research and develop cutting-edge technologies.  

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Biomedical Sciences

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School of Medicine Basic Science Building, Room 5008 http://biomedsci.ucsd.edu

All courses, faculty listings, and curricular and degree requirements described herein are subject to change or deletion without notice.

The Graduate Program

The graduate program offered by the group in Biomedical Sciences (BMS) is designed to lead students to the PhD through a combination of didactic study, laboratory rotations, and thesis research in basic and translational biomedical sciences. Research opportunities in BMS span a wide spectrum of biological and medical sciences, permitting students the options of selecting molecular, cellular, organismal, and integrated systems approaches in their research projects. Students are encouraged to design and execute original and creative research in a self-critical and independent manner. Undergraduate preparation must include courses in mathematics (through calculus), chemistry (including organic, physical, and biochemistry), and preferably participation in research. Students whose undergraduate backgrounds are significantly different will be considered provided there is sufficient evidence of interest in cell and molecular biology, genetics, pathology, physiology, pharmacology, or other disciplines in biomedical sciences, and a strong commitment to enter a field of active research and academic excellence.

Doctoral Degree Program

During the first year, the students enroll in two core courses followed by specialized courses associated with BMS training. In a required laboratory rotation program, students develop laboratory skills and the ability to formulate scientific hypotheses and become familiar with the research activities of the faculty. Students will have access to research organization in nine areas: cancer biology, computational biology and data science, genetics and genomics, cell and developmental biology, immunology, microbiome and microbial sciences, molecular and structural biology, neurobiology of disease, and molecular pharmacology and drug discovery. Students can also access training opportunities associated with organized research units led by BMS faculty such as the Glycobiology Research and Training Center (GRTC) and the Center for Academic Research and Training in Anthropogeny (CARTA). Required advanced courses and electives in subsequent years are chosen to develop students’ interests and specialized knowledge in the thesis and research areas. PhD specializations in Anthropogeny, Bioinformatics, Multiscale Biology, and Quantitative Biology, which have distinct elective requirements, are also available. BMS students are required to select their thesis advisers and begin their thesis research by the end of the first year in the program, and the average matriculation time among BMS students is between five to six years.

The graduate program is interdepartmental and interdisciplinary; it includes faculty in the Departments of Cellular and Molecular Medicine, Medicine, Pathology, Pediatrics, Pharmacology, Neurosciences, Reproductive Medicine, Chemistry and Biochemistry, Biology, Bioengineering, Psychiatry, Orthopaedics, Anesthesiology, the UC San Diego Moores Cancer Center, the Skaggs School of Pharmacy and Pharmaceutical Sciences, the Sanford-Burnham Institute, the La Jolla Institute for Allergy and Immunology, and the Salk Institute for Biological Studies.

The graduate program in biomedical sciences is also designed to educate physician-scientists and pharmacist-scientists through the School of Medicine’s Medical Scientist Training Program (MSTP) and the School of Pharmacy PharmD/PhD Program, respectively. Students already admitted to the School of Medicine or School of Pharmacy and Pharmaceutical Sciences are eligible for admission to the BMS program for doctoral training in order to receive MDs and PhDs or PharmDs and PhDs. Such students generally apply in the first or second year of their medical studies or pharmacy studies and enter graduate studies following completion of their second year. Normative time for MD and PhD and PharmD and PhD students is seven years.

Examinations

Students obtain letter grades in the core and research area courses. Candidacy for the PhD is granted following the successful completion of two research-oriented examinations. The first examination, the Research Proposition Exam, tests the student’s preparation for his or her thesis research. Preparation for the Research Proposition Exam begins as soon as students join their thesis laboratories during the first summer quarter in the program. Students prepare a written research proposal and defend the proposal in an oral examination conducted by a program-approved exam committee. The second examination is the Advancement to PhD candidacy, which takes place after the students finalize their thesis research plan, and should be completed by the end of the third summer quarter. Thereafter, the students’ thesis research progress is reviewed annually by the thesis committee. The thesis committee also approves the final dissertation. After the preparation of the dissertation, a public oral defense of the thesis completes the requirement for the PhD in the Biomedical Sciences Program.

Specialization in Immunology

A graduate specialization in immunology is offered as part of graduate training in the biomedical sciences (BMS) and School of Biological Sciences (BioSci) graduate programs. Students accepted to the biomedical sciences (BMS) graduate program will satisfy all of the degree requirements of that program. In addition, the specialization will promote an immersive educational experience in immunology that includes participation in immunology course work, seminars given by renowned speakers, biweekly research presentations by peers, a weekly journal club, and a yearly areawide immunology conference.

Admission to the Specialization and Course Requirements

Students admitted to the BMS program may enroll in the specialization in immunology after entering the program or after the first year with rotations complete. Admitted students will have satisfied the requirements for admission specified for the BMS program. Also, we expect students in graduate studies with a specialization in immunology to have completed an undergraduate course in the field or the equivalent before enrolling in the specialization.

Application Requirements

Prospective students should apply to and are admitted into the School of Biological Sciences doctoral program and can choose the specialization in immunology by notifying the immunology specialization program administrator upon admission or at any time up to completing their rotations and choosing a lab for thesis research.

The following items should be combined into a single PDF document and submitted to the program administrator ( [email protected] ):

  • Student’s CV or unofficial transcript with proof of undergraduate course work in immunology**
  • Up to one-page statement of student’s interest in interdisciplinary immunology research including career goals
  • Nomination letter from adviser acknowledging student’s academic ability and immunology interest

**For matriculation in the specialization, students just starting in the BMS graduate program should have taken the equivalent of at least one undergraduate course in immunology. Those otherwise qualified students who have not taken an undergraduate course in immunology can fulfill this requirement by taking the week-long American Association of Immunologists introductory course in immunology, presently held during the summer at the UCLA campus. Students can also fulfill this requirement by taking an upper-division summer course in immunology at any accredited college or university. Students enrolling in the specialization during or after their rotations can satisfy the requirement by completing graduate course work in immunology. The program administrator will work with admitted students to make sure they fulfill this requirement.

Specialization Requirements

  • Complete all course work and other requirements for the biomedical sciences doctorate
  • Completion of at least three approved immunology elective courses, for example: BGGN 225, Graduate Immunology, BIOM 253, Pathogens and Host Defense, BGGN 232, Innate Immunity, or Bioinformatics for Immunologists (to be added to registrar).
  • Thesis research must be completed in an approved immunology laboratory

Time Limits

Students admitted to the specialization in immunology must meet the time limit requirements as all other doctoral students in the BMS program. It is expected that students will retain the same time to degree as students not pursuing this specialization.

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Ph.D. Program

Current Students

The philosophy of the PhD program, along with the Affiliated Ph.D.  Program with the Salk Institute for Biological Studies, is to provide world-class research training in the basic biological sciences to equip a diverse group of trainees for a variety of scientific careers ranging from academia and industry to education, communication, or policy. Core principles of the program are to be student centered and attuned to the goals of the trainee.

The core curriculum focuses on development of core competencies and transferable skills in critical thinking, communication, and leadership. The first year prepares students for the core of the PhD program, the development of intellectual independence and creativity through original thesis research, guided by a thesis advisor and committee. Students have a high degree of flexibility in choice of thesis advisor through the rotation program. Throughout the program, there is strong emphasis on engaged mentoring through regular committee meetings, annual reports and Individual Development Plans.

As a central hub of the thriving San Diego biosciences community, the program maintains strong partnerships with other campus units and programs through joint faculty appointments, organized research units, and research collaborations, enabling a wide range of interdisciplinary opportunities . The mission is to conduct leading edge research in the basic biological sciences. Major areas of emphasis currently include structural biology, cell biology, developmental biology, neurobiology, immunology, microbiology, virology, plant biology, ecology, and evolutionary biology. Research in the School has emphasized studies using model organisms or in vitro mechanistic approaches, with human studies and clinical research concentrated in other departments or in the Health Sciences. Current and future areas of growth include quantitative biology, data science, and the biological consequences of climate change.

  • Rigor, reproducibility, and responsibility as hallmarks of high-quality science
  • Commitment to quality mentorships, student mental health, and well-being
  • Equity, Diversity and Inclusion as integral to program admissions and retention
  • Open science practices valuing multiple research outputs and holistic assessment of scholarly excellence
  • Communication and outreach as key aspects of scientific training

As a doctoral program embedded in a large undergraduate instructional unit, our approach incorporates substantial training in teaching methodology and best practices. Our philosophy remains that teaching and research are interdependent facets of engaged scholarship.

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Quick Links

  • Salk Institute
  • Graduate Student Handbook
  • How To Apply

Application Process

nervous cells

At the University of California, San Diego, diversity is a core component of excellence that further enhances our quality and achievement. We seek a diverse graduate student body to ensure that all of our students gain the educational benefits that result from being exposed to a broad spectrum of ideas and perspectives. These include the variety of personal experiences, values, and worldviews that arise from differences of culture and circumstance. Such differences include race, ethnicity, gender, age, religion, language, abilities/disabilities, sexual orientation, socioeconomic status, geographic region and more. We wish to broaden and deepen both the educational experience and the scholarly environment, as students and faculty learn to interact effectively with each other, preparing them to participate in an increasingly complex and pluralistic society. We also want all of our students to contribute to the campus community in a manner that enhances campus diversity and inclusiveness, consistent with the  University of California Principles of Community .

The  UC San Diego Graduate Application  can be completed on-line.

The Admissions Committee of the program considers all elements of the student's file. Top applicants are usually invited to UCSD to visit the campus and participate in faculty interviews in February and March. Arrangements are made by telephone or e-mail from the program office. It is therefore essential that you provide a correct telephone number (home and work) and email address where you can be reached during the winter and spring months. 

We recognize that the graduate school admissions process can be opaque and intimidating. To address this problem, we have established the Biomedical Application Assistance Program (BMAAP), a student-run program that aims to demystify the admissions process for applicants. For more information, please visit the BMAAP Program page.

Requirements

The  The Division of Graduate Education and Postdoctoral Affairs  at UCSD requires a minimum GPA of 3.0 for admission to graduate school. The Biomedical Sciences Admissions Committee looks specifically at an applicant’s cumulative and science GPAs, and at the types of courses taken. Recommended courses include calculus, biochemistry, organic and physical chemistry, biology, and, preferably, cell and molecular biology and mammalian physiology.

As of 2018, applicants are no longer required to submit scores for either the GRE General or Subject Tests. Applicants can optionally submit scores for the GRE General Test (verbal, quantitative, and analytical sections) and/or an applicable GRE Subject Test.  If an applicant wishes to take the GRE tests, it is advised to take the GRE in the fall prior to the fall term for which admission is sought. Applicants may self-report scores at the time of application submission. When ordering your GRE score reports, use UCSD's institution code 4836. No department codes are necessary. GRE score reports are typically received electronically within 5-7 business days from the order date. More information about the GRE may be obtained from the  Educational Testing Service  (ETS) website. 

The UCSD Graduate Application fee is $135 ($155 for international applicants). This fee may be waived for applicants who demonstrate financial hardship, US Military service, or who have participated in certain graduate preparatory programs. Please see this page for more information. For international students requesting a fee waiver, please visit this link .

Preliminary Application (International applicants)

The preliminary application is no longer required. All student applications are given equal consideration in the application review process. 

Apply for Fall 2025

Official Graduate Application Opens - early September 2024

Official Graduate Application Deadline - approximately late November 2024

Interviews and Recruitment (In-Person) Weekends - to be announced, most likely in February 2025

  Begin Your Application

More information.

Frequently Asked Questions

Please contact the Biomedical Sciences Graduate Program offfice for more information or questions: [email protected].

Statement of Purpose

Focus your Statement of Purpose on the reasons you are interested in attending the UC San Diego Biomedical Sciences Graduate Program. The statement has a 1500 word limit and should be well organized, concise, and completely free of grammar, punctuation, and spelling errors.

Include responses to the following as part of your statement:   

  • Summarize the long term goals of research in which you participated.
  • Describe one research problem, project or area for graduate study that excites you. Have you thought about what you want to accomplish in graduate school and how is UC San Diego a good match for these goals?

Letters of Recommendation

These are of great value in assessing a student’s qualifications for a career in research. It is advantageous to have letters submitted by well-known faculty who can critically evaluate a student’s academic performance, undergraduate research experience, and potential for a career in biomedical sciences.

Research Experience

Undergraduate and/or post-college research experience is considered an important part of a student’s preparation for graduate work. It gives the student the opportunity to experience laboratory work and thus make a knowledgeable decision to pursue a career in basic biomedical sciences.

Official Application

Screening of applications will begin in late November. It is recommended that applicants submit their application well before the deadline to be considered for on-campus interviews. Typical causes of delayed consideration include missing letters of recommendation and transcripts. 

PLEASE NOTE: To expedite the processing of your application, the Biomedical Sciences Admissions Committee requires that you upload PDF versions of your official or unofficial transcripts directly into the UCSD on-line application system. More information on this process is available in the on-line application system.

Non-discrimination Statement

The University of California, in compliance with Title VI of the Civil Rights Act of 1964, Title IX of the Education Amendments of 1972, Section 504 of the Rehabilitation Act of 1973, and the Age Discrimination Act of 1975, and the Americans with Disabilities Act of 1990, does not discriminate on the basis of race, color, national origin, religion, sex, disability, or age in any of its policies, procedures, or practices; nor does the university discriminate on the basis of sexual orientation. This nondiscrimination policy covers admission and access to, and treatment and employment in, university programs and activities, including but not limited to, academic admission, financial aid, educational services, and student employment.

Inquiries regarding the university's equal opportunity policies may be directed to the campus compliance coordinator, (858) 534-0195.

Biomedical Sciences

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Doctoral Programs

Department doctoral programs.

The School of Biological Sciences offers graduate training in a collaborative environment with several schools and graduate programs. This provides excellent research training and professional development for graduate students pursuing the doctorate. Training opportunities include broad areas of Biological, Biomedical and Environmental Sciences including Neuroscience, Immunology and Pathogenesis, Cancer Biology, Developmental and Stem Cell Biology, Systems and Computational Biology, Genomics, Structural Biology, Ecology, and Evolutionary Biology.

Ecology and Evolutionary Biology, PhD

Research in Ecology and Evolutionary Biology uses a range of methodologies spanning field studies, experimental approaches, and computational approaches. Students and faculty have easy access to field sites that include marine, desert, and California chaparral communities. The faculty have diverse interests including evolutionary biology, population genetics, genomics, behavioral ecology, physiology, and global change biology. Faculty in Ecology and Evolutionary Biology have extensive collaborations with scientists in neuroscience, molecular biology, genomics and microbiomes, global climate biology, systems biology, mathematics, as well as with faculty at other institutions.

The Department of Ecology and Evolutionary Biology offers admission directly to a departmental graduate program with full financial support. Students initiate research with a faculty thesis advisor while they take required coursework and complete other degree requirements. 

Learning Outcomes by Department

The Department of Ecology and Evolutionary Biology is the only department that offers direct admission to a PhD Program. The other Dunlop Dunlop School departments offer admission to their PhD programs through a gateway disciplinary program such as CMB or INP. Click below to learn more about learning outcomes by department.

Neurobiology and Behavior, PhD

Charlie Dunlop School of Biological Sciences   PLOs

Neurobiology and Behavior M.S./Ph.D.

PLO1: Core Knowledge

  • Demonstrate a basic knowledge of central concepts in the biological sciences
  • Understand the current concepts in molecular biology, biochemistry, and biomedical sciences
  • Demonstrate specialized knowledge of cellular and molecular biology sufficient to carry out substantive independent research

PLO2: Research Methods and Analysis

  • Read and critically evaluate the scientific literature
  • Formulate hypotheses based on current concepts in the field
  • Design, conduct, and interpret their independent research projects
  • Understand the range of tools appropriate for research in their sub-field
  • Understand and follow research ethics

PLO3: Pedagogy:

  • Communicate effectively to large and small groups in pedagogical settings including teaching, research seminars, and other formats
  • Identify and effectively deploy suitable technologies for use in all aspects of instruction

PLO4: Scholarly Communication:

  • Review and discuss relevant literature and their significance
  • Publish research results in peer-reviewed publications and in a dissertation
  • Communicate research results effectively through oral presentations at scientific seminars, conferences, and other venues
  • Make clear and cogent oral presentations, including effective use of technology

PLO5: Professionalism

  • Write compelling abstracts describing their research for consideration at research conferences
  • Prepare oral presentations suitable for presentation at a research conference
  • Make effective contributions to research teams and learning seminars
  • Make effective contributions to department, university, community, and professional service
  • Mentor junior researchers (e.g., undergraduates, beginning graduate students)

PLO6: Independent Research

  • Develop their own research projects that meet high standards of theoretical and methodological rigor with lasting impact
  • Produce scholarship that is comparable in scope and format to articles that appear in leading peer-reviewed journals in molecular and biomedical sciences
  • Supervise junior researchers (e.g., high school students, undergraduates, beginning graduate students) effectively

Molecular Biology and Biochemistry, PhD

Molecular Biology and Biochemistry M.S./Ph.D.

  • Master current concepts in molecular biology, biochemistry, and biomedical sciences
  • Acquire specialized knowledge of cellular and molecular biology sufficient to carry out substantive independent research

PLO2: Research Methods and Analysis 

  • Design, conduct, and interpret experiments to complete an original research project
  • Understand the range of tools appropriate for research in the specific sub-field
  • Appreciate and adhere to research ethics

PLO3: Pedagogy: 

  • Communicate effectively to small and large groups in pedagogical settings such as teaching and research seminars

PLO4: Scholarly Communication: 

  • Review and discuss relevant literature and its significance
  • Complete an individual development plan (IDP) at the time of entering the MBB program (Fall quarter of the 2 nd year) and update it annually
  • Submit fellowship proposals to private and governmental agencies to solicit independent funding for graduate research
  • Anticipate and meet the needs for professional transitions in a timely fashion (prior to degree completion)
  • Write compelling abstracts describing research for consideration at research conferences

PLO6: Independent Research 

  • Effectively supervise junior researchers (e.g., high school students, undergraduates, beginning graduate students)

Interdepartmental Doctoral Gateway Programs

Gateway programs offer admission to the doctoral programs affiliated with the Charlie Dunlop School of Biological Sciences, School of Medicine, School of Physical Sciences, Engineering and Information & Computer Science. Students enroll for the first academic year while they do lab rotations and take required coursework. Then, students select a thesis advisor and transfer to a department and complete remaining degree requirements. Gateway programs offer students excellent opportunities to perform laboratory rotations with any of a large number of faculty participants in that program, and in many areas of biological sciences.

Cellular Molecular Dunlop Schoolences

The PhD program in Cellular & Molecular Biosciences (CMB) at UC Irvine provides ideal training to launch the careers of talented researchers in diverse fields of biological and biomedical sciences. With five different study focuses, the CMB PhD program gears future scientists to be ready for a diverse field.  The program offers a rigorous but flexible curriculum with an extensive choice of laboratories and allows students to tailor their training to individual interests and goals. Outstanding facilities, a collaborative culture, a commitment to diversity, and guaranteed on-campus housing all contribute to a productive graduate experience. 

Interdepartamental Neuroscience Program

The Interdepartmental Neuroscience Program (INP) provides a vehicle for meeting the diversity and challenges of graduate training in such a broad discipline. Neuroscience is an inherently broad and multidisciplinary area of scientific pursuit and scholarship. It has intellectual links to fields as diverse as developmental and cell biology, molecular biology, physiology, pharmacology, anatomy, psychology, computer science, and physics. The substantial breadth of Neuroscience is one of its strengths as a discipline, and one of the features that makes it an attractive and important area for graduate study. Students may train with any participating faculty member and are exposed to a variety of approaches before deciding on a research area for focused dissertation work. After the initial year of academic coursework and laboratory rotations, students join the more specialized graduate program of their chosen thesis advisor.

Mathematical, Computational, and System Biology

The goal of UCI’s program in Mathematical, Computational and Systems Biology (MCSB) is to provide students from a variety of academic backgrounds with doctoral training suitable for research careers in the nascent field of Systems Biology. The program emphasizes in-depth classroom study, interdisciplinary research rotations, and individualized advising. The MCSB Program is supported by funding from UCI’s Graduate Division, by a National Institute of General Medical Sciences grant to the UCI Center for Complex Biological Systems, and an NIH Training Grant.

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Program in Biomedical Sciences & Engineering

Grad Student

The Graduate Program in Biomedical Sciences and Engineering (PBSE) reflects the interdisciplinary and collaborative nature of biomedical research at UCSC. PBSE predoctoral fellows have the opportunity to study biology at the molecular, cellular, and systems levels, and can take advantage of advanced laboratory facilities, computational tools, and a highly collaborative research environment. First-year students can do research rotations with over sixty faculty members, who come primarily from the Departments of Molecular, Cell, and Developmental Biology, Biomolecular Engineering, Chemistry and Biochemistry, and Microbiology and Environmental Toxicology. Students apply directly to the PBSE, selecting from the following four interdisciplinary training tracks:

  • Biomolecular Engineering and Bioinformatics (BMEB)
  • Chemical Biology, Biochemistry and Biophysics (CB3)
  • Microbiology and Environmental Toxicology (METX)
  • Molecular, Cell and Developmental Biology (MCD)

Each training track offers a complete set of core courses, electives, seminars and journal clubs that provide rigorous training in each of the four broad areas. Students may switch training tracks, and we encourage students to take advantage of PBSE's unique opportunities for carrying out interdisciplinary research.  A designated emphasis in Stem Cell Research will soon be available for all tracks. Questions regarding PBSE may be sent to Dr. Angela Brooks , Director of the program.

Welcome from PBSE Director Dr. Angela Brooks

PBSE is a unique Ph.D. program at UC Santa Cruz designed to facilitate interdisciplinary graduate training in the biomedical sciences. Our students have access to world-class facilities and collaborate with experts in a variety of fields. They make significant contributions to research and go on to pursue diverse careers in biology, medicine, and engineering. I invite you to explore our web pages and contact us with any questions.

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Last modified: June 21, 2023 128.114.113.87

biomedical science phd california

Best Biomedical Sciences colleges in California 2024

Best biomedical sciences colleges in california for 2024.

biomedical science phd california

University of California-San Diego offers 2 Biomedical Sciences, General degree programs. It's a very large, public, four-year university in a large city. In 2022, 31 Biomedical Sciences, General students graduated with students earning 30 Doctoral degrees, and 1 Master's degree.

biomedical science phd california

University of California-Davis offers 1 Biomedical Sciences, General degree programs. It's a very large, public, four-year university in a small suburb. In 2022, 16 Biomedical Sciences, General students graduated with students earning 16 Bachelor's degrees.

biomedical science phd california

University of California-Santa Cruz offers 1 Biomedical Sciences, General degree programs. It's a large, public, four-year university in a small city. In 2022, 62 Biomedical Sciences, General students graduated with students earning 62 Bachelor's degrees.

biomedical science phd california

University of Southern California offers 2 Biomedical Sciences, General degree programs. It's a very large, private not-for-profit, four-year university in a large city. In 2022, 6 Biomedical Sciences, General students graduated with students earning 5 Doctoral degrees, and 1 Master's degree.

biomedical science phd california

University of California-Riverside offers 2 Biomedical Sciences, General degree programs. It's a very large, public, four-year university in a large city. In 2022, 14 Biomedical Sciences, General students graduated with students earning 8 Doctoral degrees, and 6 Master's degrees.

biomedical science phd california

San Francisco State University offers 2 Biomedical Sciences, General degree programs. It's a very large, public, four-year university in a large city. In 2022, 11 Biomedical Sciences, General students graduated with students earning 8 Master's degrees, and 3 Certificates.

biomedical science phd california

Stanford University offers 1 Biomedical Sciences, General degree programs. It's a large, private not-for-profit, four-year university in a large suburb. In 2022, 8 Biomedical Sciences, General students graduated with students earning 8 Master's degrees.

biomedical science phd california

Biola University offers 1 Biomedical Sciences, General degree programs. It's a medium sized, private not-for-profit, four-year university in a large suburb. In 2022, 21 Biomedical Sciences, General students graduated with students earning 21 Bachelor's degrees.

biomedical science phd california

Mount Saint Mary's University offers 1 Biomedical Sciences, General degree programs. It's a small, private not-for-profit, four-year university in a large city. In 2022, 3 Biomedical Sciences, General students graduated with students earning 3 Bachelor's degrees.

biomedical science phd california

Loyola Marymount University offers 1 Biomedical Sciences, General degree programs. It's a large, private not-for-profit, four-year university in a large city.

Find local colleges with Biomedical Sciences majors in California

List of all biomedical sciences colleges in california.

School Average Tuition Student Teacher Ratio Enrolled Students
La Jolla, CA 3/5 30 : 1 42,006
Davis, CA 3/5 23 : 1 39,679
Santa Cruz, CA 3/5 25 : 1 19,478
Los Angeles, CA 5/5 23 : 1 48,945
Riverside, CA 3/5 29 : 1 26,809

Program in Biomedical Science and Engineering: Molecular, Cell and Developmental Biology

The Program in Biomedical Science and Engineering: Molecular, Cell and Developmental Biology (PBSE-MCD) track offers a Doctoral degree. 

Important: Transcripts and Letters of Recommendation are required of all applicants. 

Read the  application instructions ​ page for full details., see the test score requirements for this program., below are specific requirements also needed for this program:.

Résumé/CV (required)

Statement of Purpose (required):

​What would you like to study in graduate school and how does research in Molecular, Cell, and Developmental Biology (MCDB) Department align with your scientific interests? What are your academic and career goals? Limit your answer to one page. 

A Statement of Previous Research (required):

Please describe your research experience: describe the motivations behind one (or more) of your research projects, including experimental tests of theories or hypotheses. What have you learned from this experience? It is more important that you address the points above for one project in depth, rather than to provide a comprehensive list of all of your research experiences. In any case, limit your statement to one page.

A Personal History Statement (required):

Describe any educational, familial, health, cultural, economic, or social challenges you may have faced. Limit your statement to one page. 

See the application deadline

Contact the program

Read more about this program:

https://pbse.ucsc.edu/mcd/index.html

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Biomedical Sciences

Biomedical Sciences from University of California, San Francisco  is an interdisciplinary graduate research program that equips students with the training and research tools to dissect disease-related biology, from single cells to tissue and organ systems. 

University of California, San Francisco Multiple locations San Francisco , California , United States Top 4% worldwide Studyportals University Meta Ranking 3.0 Read 2 reviews

Biomedical Sciences from University of California, San Francisco curriculum results in a new generation of interdisciplinary biomedical scientists who are able to forge collaborations that break down traditional research boundaries.

Students in the Biomedical Sciences program from University of California, San Francisco will:

  • acquire a level of competence in molecular biology, genetics, and cell biology comparable to that expected of students in traditional programs focused in these areas. 
  • get a rigorous and molecular study of core developmental, physiological, and pathological features of human biology and disease.

Programme Structure

Courses include:

  • Cancer Biology and Cell Signalling
  • Human Genetics
  • Neurobiology
  • Vascular and Cardiac Biology
  • Cell Biology
  • Investigating Human Biology and Disease 

Key information

  • 60 months

Start dates & application deadlines

  • Starting 2024-08-26 00:00:00 Application deadline not specified.

Disciplines

Academic requirements, english requirements, student insurance.

Make sure to cover your health, travel, and stay while studying abroad. Even global coverages can miss important items, so make sure your student insurance ticks all the following:

  • Additional medical costs (i.e. dental)
  • Repatriation, if something happens to you or your family
  • Home contents and baggage

We partnered with Aon to provide you with the best affordable student insurance, for a carefree experience away from home.

Starting from €0.53/day, free cancellation any time.

Remember, countries and universities may have specific insurance requirements. To learn more about how student insurance work at University of California, San Francisco and/or in United States, please visit Student Insurance Portal .

Other requirements

General requirements.

  • A minimum of a bachelor's degree (BA/BS) or the equivalent from an accredited institution is required. (Some programs or pathways may require a master’s degree. Please check the program that you are interested in to confirm the application requirements.)
  • To be eligible for admission, you must have at least a 3.0 (B) grade point average.
  • You may apply to only one graduate program per application cycle/year.
  • Unofficial transcripts may be submitted for review purposes only. Final official transcripts must be submitted upon matriculation if an offer of admission is made.
  • Upon completion of the application and receipt of the non-refundable application fee

Tuition Fee

International, living costs for san francisco.

The living costs include the total expenses per month, covering accommodation, public transportation, utilities (electricity, internet), books and groceries.

In order for us to give you accurate scholarship information, we ask that you please confirm a few details and create an account with us.

Scholarships Information

Below you will find PhD's scholarship opportunities for Biomedical Sciences.

Available Scholarships

You are eligible to apply for these scholarships but a selection process will still be applied by the provider.

Read more about eligibility

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School of Medicine News

A Community-Based Medical School for the 21st Century

Dr. Zheng with members of his lab

RNA and the Cycle of Life

Sika Zheng, PhD has helped UCR become a hub for RNA research with the potential to help treat neurodegenerative diseases.

As a child, Sika Zheng, PhD, founding director of the Center for RNA Biology and Medicine and a professor of biomedical sciences at the UC Riverside School of Medicine , was fascinated by the cycle of life and death. While he initially planned to become a physician, his curiosity about the process from birth to decay and the workings of the brain led him to pursue a PhD in neuroscience at Johns Hopkins University with a focus on neurodegeneration.

“In the lab at that time, most people were trying to study how neurons died,” Zheng recalled. Yet rather than focusing on neurodegeneration in conditions like Parkinson’s disease and stroke, he continued, “I started thinking about how you can make neurons live longer and finding protective mechanisms.”

Tiny cells, significant impact

This focus on neuronal longevity became the basis for Zheng’s career. While there’s constant turnover of other cells in the body, the same neurons remain with each person throughout their lifespan. Zheng dove into examining neurons at the molecular level during his postdoctoral research at UCLA and began investigations into alternative RNA splicing, essentially the cellular process that allows a gene to produce multiple RNA molecules of different sequences and functions in different cells. (RNA is the copy of a cell’s DNA that translates genetic code to create proteins and drives other cell processes.)

Sika Zheng, PhD

“At the time, little was known about the functional significance  of alternative splicing even though this process had been studied in great detail at a biochemistry level,” Zheng said. “Very rarely was it being investigated in a system, in a live animal, so that's where I joined in.”

Zheng focused on bridging the fields of RNA and neuroscience to make new discoveries, an objective he continued to pursue when he joined the UC Riverside School of Medicine in 2014. “Before I started, the RNA field and neuroscience field didn't really talk to each other,” he explained. Bringing them together allowed him to investigate unique RNA regulations in the brain and, conversely, unveil the specific contributions of neural specific splicing to brain biology.

In Zheng’s lab at UCR , he has investigated neuronal longevity by looking into aspects including the impact of neurological diseases on RNA and the effects of specific gene splicing on cell death pathways, including apoptosis, or programmed cell death. His research in this area may benefit health issues in the community and beyond, building on the UCR SOM’s commitment to improve health in the region.

“Our research has a broad impact, first for fundamental knowledge but also potentially for neurological and neurodegenerative disease,” Zheng said. “Some of these diseases are quite overrepresented in the area. So even though we may not be immediately making therapeutic interventions or designing therapeutic strategies, our research is highly relevant.”

He has also shared his research results with the community, recently visiting the UCR Palm Desert Center to give a talk on his findings and connect with residents. “From feedback we got, it was pretty well received, and there was a lot of interest in our discoveries,” he said. “More broadly speaking, I think we're all trying to increase UCR’s visibility and by doing that, enhance UCR's capacity to serve the Inland Empire better.”

Putting UCR on the map with a new RNA center

Conducting groundbreaking research while promoting UCR are also goals behind a new interdisciplinary center Zheng founded, the UCR Center for RNA Biology and Medicine . The center includes three dozen labs from four colleges across the campus: the School of Medicine (SOM) , the College of Natural & Agricultural Sciences (CNAS) , the Marlan and Rosemary Bourns College of Engineering (BCOE) , and the College of Humanities, Arts, and Social Sciences (CHASS) .

“We want this center to be the innovation hub of UCR as we bring people together and create science that individual labs would not necessarily be able to generate, and through that train lots of next-generation scientists,” said Zheng, who serves as the center’s director. “I've been putting in lots of effort with my RNA colleagues to jumpstart the center and make it happen, and feel like it is a new baby for my journey,” he added. “I really look forward to its success.”

The center began with an inaugural RNA Symposium in November 2022 . Adrian R. Krainer, PhD, who won the 2019 Life Sciences Breakthrough Prize, presented the keynote lecture, with speakers from universities across California in attendance. Zheng said he hopes to make the symposium an annual event, but said the center has already succeeded in raising UCR’s profile.

“People started realizing we are UC Riverside, not UC Irvine,” he recalled. “Now, they know that we have an RNA center here at UC Riverside.”

The center brings together RNA researchers from across UCR’s campus to combine, organize, and strengthen their efforts. “We provide a platform… to get together to talk about research on RNA and exchange ideas,” Zheng said. While a couple of RNA researchers had previously tried to organize smaller initiatives to facilitate working together, Zheng said the lack of funding and being spread out among various departments made it unsustainable.

“Now, I am using the center’s funds to kickstart something to put many more research groups together,” Zheng said. “It's exciting because we want to build something new.”

Zheng has already seen the center make a difference on campus as well. Several UCR researchers who had never studied RNA biology had their first grants on RNA research awarded at the end of 2023 and the beginning of 2024. “Those are examples that the center is promoting individual investigators directly and then also helping the university to boost their visibility,” Zheng said.

Mentoring the “true heroes” of science

Zheng’s dedication to his research is clear, but so is his commitment to his students and trainees.

Zheng said he enjoys working with students and postdocs both in and out of the lab. “They are the true heroes behind all of these scientific discoveries,” he pointed out, noting that students and postdocs are often the ones handling pipettes or taking care of animals in the lab.

“It's great to see that your advice and guidance can have some positive influence,” Zheng added. “To see how they transform with your input over time--I really cherish those moments, particularly when some of them work with you for many years.”

David Nikom, now a doctoral candidate in his fourth year, joined Zheng’s lab during his first year of graduate school. Nikom recalled Zheng encouraging him to develop a class paper on alternative RNA splicing in disease into a textbook chapter. “After sending him my revised draft he said it deserved to be in a journal, rather than [as a] book chapter, which felt amazing,” Nikom said. They published the review paper together in Nature Reviews Neuroscience in 2023 .

“Dr. Zheng sets the bar high because he knows you can reach it,” Nikom said. “It’s easier to work hard when your mentor leads by example, and Dr. Zheng embodies a work ethic and discipline that I hope to achieve in my own research career.”

Zheng said he learns from his students, too, such as former graduate student John Vuong, PhD. "John spearheaded developing a new sophisticated lab technique in the absence of local expertise," said Zheng. He added, however, that Vuong was misunderstood by some individuals who didn’t realize that his late arrivals to work reflected long nights in the lab rather than a lack of dedication to his work. Similarly, while Vuong’s research ideas were strong, Zheng said his initially modest presentation skills contributed to him getting some negative feedback from his thesis committee.

John Vuong, PhD with several UCR friends

“John taught me that we should give students some freedom to explore,” Zheng said of Vuong, who went on to become a scientist at the biotech company Thermo Fisher Scientific. “At a certain point you feel like, this is going to be a student who needs a lot of hand-holding in one way or another,” he continued. “But it turns out that if you just let him be himself, he knows what he's doing. With some coaching and mentoring, like teaching him how to do a presentation, he will shine.”

For his part, Vuong recalled Zheng’s evident excitement when Vuong made a breakthrough in a project that later became his dissertation. "As a researcher, Sika was at times demanding but helped instill values of perseverance, accountability, and attention to detail," Vuong said. "Sika was supportive in my time as a budding researcher in providing key insights and lessons leading to breakthroughs in my dissertation and was instrumental as a mentor helping me earn my PhD."

In the end, Zheng said, the goal is to do good science while simultaneously setting each student up for the future with a solid scientific background. “Most importantly, when experiments fail, we don't want a student to fail,” Zheng said. “We want to train them in a way that even if the experiment fails, they can still be a scientist or at least have a scientist's mind with critical and independent thinking.”

This remains true, he added, even if the student ends up pursuing a different career path. “It's our job to really train their scientific mind, for them to analyze and understand things in a scientific way,” he said. “Even if they don't necessarily stay in academia, if they're equipped with that kind of mindset, I think they're going to do good things in the world.”

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UT Austin Biomedical Engineering Graduate Program Among Top 20 in U.S.

UT Austin Biomedical Engineering Student In Lab

The University of Texas at Austin Department of Biomedical Engineering's graduate program ranked No. 17 in U.S. News & World Report’s 2024-2025 graduate engineering program rankings, released on Tuesday.

Graduate students have a variety of options to choose from. Many students take advantage of a 100% fully-funded Ph.D. with coursework that includes a Personal Learning Plan—tailoring educational experiences to meet individual needs.

Students can also choose from a  B.S.E./M.S.E. Integrated degree . This program allows students to earn both a Bachelor of Science degree in biomedical engineering and Master of Science degree in engineering in five years.

A student's coursework is selected in consultation with the student's supervisor, dissertation committee and the graduate advisor to ensure adequate breadth and depth of knowledge across our eight research areas .

Lastly, there is a M.D./M.S. Dual Degree Program. In collaboration with Dell Medical School , we offer an M.S. degree specifically designed for current medical students. Dell Medical students can choose the biomedical engineering M.S. to fulfill requirements in year 3, also known as the Growth Year. The overarching goal of the program is to produce physicians who can practice medicine and biomedical research with the quantitative analytical skills of an engineer.

The Cockrell School of Engineering once again finished among the top 10 engineering programs in the nation, coming in at No. 8. In addition, three programs ranked in the top five, seven in the top 10 and all programs finished in the top 20.

U.S. News & World Report’s annual engineering graduate program rankings are based on student acceptance rates, faculty/student ratios, research expenditures, reputation among engineers in industry and academia, the number of doctoral degrees produced, the percentage of faculty elected to the National Academy of Engineering and entrance exam scores.

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  • PhD applications in Biomedical Science and Biochemistry

biomedical science phd california

Are you a high-achieving student interested in pursuing a PhD in molecular and biomedical sciences? Do you want to work with top researchers, in a world-class research facility, in a beautiful city at the bottom of the world? Then check out the Division of Biomedical Science and Biochemistry at the Australian National University.

Our Science : We provide an outstanding research environment with a world-class reputation. We are an interdisciplinary division, with highly integrated labs that work at the forefront of membrane transport processes, host-pathogen biology, drug and vaccine development, immune processes, cancer biology and evolutionary genomics. We research a wide range of topics – from the physiology of intracellular parasites to the assembly of bacterial nanomachines; from the structure and biophysics of membrane transporters to the early evolution of animals. If you are interested in doing a PhD with us check out our Research Groups below, and contact prospective supervisors.

  • Adamska Group - Genomic and evolutionary basis of animal development
  • Altin Group - Tumour immunology and liposome targeting group
  • Behm Group - Nematode molecular biology
  • Brock Group - Membrane structural biology, biochemistry and biophysics
  • Broer Group - Membrane transport and nutrition
  • Casarotto Group - Biomolecular interactions  
  • Corry Group - Transport proteins and computational biophysics
  • Fahrer Group - Immunology
  • Furlong Group - Bacterial structural defence
  • Jackson Group - SynBio
  • Lehane Group - Antimalarial drug action and resistance
  • Leyton Group - Assembly, function, and applications of nanoscale bacterial surface structures
  • Maier Group - Molecular mechanisms of malaria pathogenesis
  • Saliba Group - Physiology and biochemistry of the malaria parasite
  • Spry Group - Drug discovery for infectious diseases
  • Tham Group - Understanding chytrid infection biology
  • van Dooren Group - Cell biology and metabolism of apicomplexan parasites
  • Verma Group - Bacterial and bacteriophage genetics, and vaccine development.  

Our Values : Our division includes award-winning supervisors and teachers, and we pride ourselves on providing high-quality supervision to our students. Our PhD students are well supported through internal funding, including for conference travel and computational resources, and our labs are located in a well-equipped, modern research facility. We have a thriving community of PhD students, and our graduates go on to productive careers in many areas of science and beyond. We value diversity and inclusivity in science and have active policies to prevent discrimination. Our faculty and their research teams are comprised of people from all over the globe and from all walks of life.

Location : The ANU is situated in Canberra, Australia’s capital city. Canberra is a well-resourced regional city, set amongst beautiful mountains and surrounded by eucalypt forests. Bike riding and hiking are everyday activities, and we are a two-hour drive away from both winter snowfields and some of the most beautiful beaches on Earth. Canberra’s birdlife is stunning, and we regularly have to brake for kangaroos on our daily commute. Despite its regional setting, Canberra is a vibrant, multicultural city. There are frequent cultural evenings, festivals, art exhibits, music events, and world-class restaurant and coffee scenes.

The Australian National University: We are a research-intensive university and have an international reputation for research excellence. The ANU is frequently ranked as Australia’s top university, and one of the top 50 universities in the world.

Candidate eligibility : Domestic applicants must be Australian citizens or permanent residents, or New Zealand citizens, and must have previously completed a 1 st  class Honours or Masters thesis in a relevant field. Fully funded International PhD scholarships are highly competitive at the ANU. To be a viable candidate you need: (1) excellent undergraduate marks; (2) a Masters by Research with a high mark; or (rarely) equivalent proof of research experience. (3) While not essential, having published research papers in the past is an advantage.

What do I do? First, contact potential supervisors from the list of staff on the Research Groups above (include your CV and grades, and some details about what it is that attracts you to their field of research). If they are interested in supervising you, you can submit an application to commence a PhD via the online link below. Your application should include a detailed CV, full academic transcripts, the name of your potential supervisor, and a brief description of a research project that you and your potential supervisor have discussed. Entry to the PhD program is open to applicants with a Bachelor degree that have also completed (or are in the process of completing) an Honours or Masters research degree that comprises at least a half year, full time research component and a thesis (5,000-10,000 words). Applicants with significant research experience and publications may be deemed eligible if their achievements can be justified as equivalent to completing an Honours/Masters degree. As part of your application for undertaking a PhD, you will automatically be considered for a stipend scholarship if you tick the scholarship box in the application. Your potential supervisor or the Biomedical Science and Biochemistry graduate convenor, Dr Giel van Dooren, can advise you on how competitive your application is likely to be for obtaining a stipend scholarship. If you have questions about any aspect of the PhD application process, you can contact Giel van Dooren ( [email protected] )

The application form is here , along with general information on how to apply and the details about the Doctor of Philosophy program . PhD applications are due by 15 April (the mid-year round for both international and domestic students),  31 August (international students), or 31 October (domestic students). Your application will require:

  • Academic transcripts and graduation/completion certificates from your Bachelor and Honours/MSc degree (in its original language and, if needed, a translation), with grading scale information.
  • A current CV, including scholarships or prizes, publications and conference presentations.
  • A research proposal (this is quite short and your potential supervisor can help you with it).
  • Proficiency in English: For international students from many countries, a current IELTS or TOEFL certificate is required. 
  • Three referee reports. You need to enter your referee details on the application form. The system then automatically sends them a link to complete an online form. NB: You need to ensure your referees are willing and able to write recommendation letters and do so promptly. Plan ahead and alert them.

* If you are an international student and have the possibility of PhD funding from your home country or elsewhere, let a potential supervisor know.

Morningside Graduate School of Biomedical Sciences celebrates milestones of IMSD, PREP students

By Pat Sargent and Amelia Rodriguez

Group shot of PREP students

The Morningside Graduate School of Biomedical Sciences at UMass Chan Medical School hosted its year-end celebration to highlight a successful academic year and the 22 members of the Postbaccalaureate Research Education Program (PREP) and the Initiative for Maximizing Student Development (IMSD) program.

The event recognized students who completed their PhDs, are finishing PREP or have successfully passed their PhD qualifying exam.

The PREP and the IMSD program are funded through the National Institutes of Health with the shared goal of increasing diversity and promoting the success of students in biomedical sciences. 

Group shot of the members of the 2024 PREP class

Enhancing academic preparedness and research experience Joshuel Velasco has been enrolled in the PREP program since 2022, working in the lab of Silvia Corvera, MD, the Endowed Chair in Diabetes Research  and professor of molecular medicine. Velasco will be pursuing his PhD in biomedical sciences at UMass Chan this fall.

“Before starting PREP, I wasn’t sure that I was capable of being a PhD student in biomedical sciences, but with the experience that I’ve gained, I am definitely confident that I can succeed,” Velasco said. “I wanted to hone my scientific confidence. I feel I accomplished exactly that. I was knee deep into the research, doing my own project, and having the mentors along the way to steer me in the right direction.”

“The IMSD program helped me understand what being a research-focused scientist means . . . I’m very glad I’ve been a part of this community.”

The PREP program runs from July to June and enrolls nine students each year. To be eligible, students must be U.S. citizens or permanent residents and have completed a bachelor’s degree within the past three years and intend to pursue doctoral study in the biomedical sciences.

PREP student Amanda Modica will be pursuing a PhD in microbiology and immunology at Drexel University College of Medicine Graduate School of Biomedical Sciences and Professional Studies starting this fall.

During her PREP year at UMass Chan, Modica worked in the lab of Kiera Clayton, PhD, assistant professor of pathology, investigating HIV accessory proteins and the mechanisms by which they trigger natural killer cells.

“It was a fantastic experience. I gained the skills I needed to bolster my graduate school application and I feel well prepared for the next step,” Modica said.

Building a community of underrepresented students in the biomedical sciences David Upegui, PhD, a science teacher at Central Falls High School in Rhode Island and an adjunct professor of education at Brown University, served as the year-end celebration’s keynote speaker. Coincidentally, Helen Magana, an IMSD student and fourth-year biomedical science PhD candidate, had Dr. Upegui as her science teacher at Central Falls High.

Photo of Kate Lapane, Whitney Bentley, Brian Lewis and Zhané Adamson inside a conference room

“The IMSD program helped me understand what being a research-focused scientist means,” said Magana, who has a Bachelor of Science in biology from Rhode Island College. “I think of all the supplementary workshops that we get—writing, presenting, how to present data. How we think about those kinds of things isn’t anything I really thought about until I was with IMSD.”

Magana works in the lab of Mary Munson, PhD, professor of biochemistry & molecular biotechnology, vice chair of diversity for the department, and assistant vice provost for health equity, where she researches specific mutations in the VPS45 gene that causes congenital neutropenia, a rare, severe disorder in newborns.

The IMSD program enrolls more than 40 students at all stages in their PhD journey. The program strives to increase the number of students who complete PhD degrees despite socially, culturally, economically or educationally disadvantaged backgrounds that inhibit entry into research training and careers.

“Everyone in the program comes from different backgrounds, it’s very collaborative, and UMass Chan really wants to see us succeed in the program,” said Calvin Johnson, a PhD candidate in the IMSD program who earned his Bachelor of Science in microbiology from UMass Amherst. He works in the lab of Jason Pitarresi, PhD, assistant professor of medicine, where he is identifying new drug targets for reversing suppression of the immune system in pancreatic cancer.

“The faculty have been supportive, friendly and eager to answer questions for trainees.”

IMSD Class of 2024 Nathan Bamidele Xavier Gonzalez Leslie Torres Ulloa

PREP Class of 2024 Lali Berelashvili Tarrin D’Aniello Amanda Modica Caroline Moore Blanca Muñoz Villarreal Rachell Rivera Olachi Unaka Joshuel Velasco Astrid Veloz-Maury

IMSD Qualifying Exam Completion Zhané Adamson Rakeyah Ahsan Brianna Dominguez Calvin Johnson Oscar Lam Rebecca Pavchinskiy Claudia Perez Alejandra Rivera Nieves Carolyn Senneca Askar Temirbek

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Doreen Joseph Receives Inaugural Esogbue-May Award in Engineering Diversity

  • by Molly Bechtel
  • June 24, 2024

Doreen Joseph, a Ph.D. candidate in computer science at the University of California, Davis, is the inaugural recipient of the Professor Emeritus Augustine O. Esogbue and Chancellor Gary May Endowed Award in Engineering Diversity.

Doreen Joseph

The award was established in 2023 with a gift from Augustine O. Esogbue , professor emeritus in the H. Milton Stewart School of Industrial and Systems Engineering at Georgia Institute of Technology and a longtime mentor to Chancellor Gary S. May. It recognizes graduate students who share a passion for excellence and demonstrated commitment to advancing diversity, equity and inclusion in the field of engineering and computer science, especially through dedication to enhancing UC Davis' recruiting, retaining and graduating Black Ph.D. students in STEM fields.

Before joining UC Davis, Joseph attended George Mason University, where she was the first Black woman to earn a Bachelor of Science degree in cyber security engineering. Her research focuses on using machine learning as a tool to analyze and develop more secure firmware for embedded devices, such as phones.

Several mentors, including Renetta Garrison Tull, vice chancellor of diversity, equity and inclusion and adjunct professor in the Department of Electrical and Computer Engineering, encouraged Joseph to attend UC Davis. Joseph and Tull met at the convention of the National Society of Black Engineers, the same organization that led to the decades-long relationship between Esogbue and May.

Joseph has been a steadfast advocate for DEI at UC Davis. Informally, she builds community among Black Ph.D. students through game nights at her house. Formally, she has contributed to DEI initiatives in the computer science department, on campus and at the University of California systemwide level. These include the Computer Science Program-level Assessment Capacity on Enrichment for Equity Committee , the Chancellor's Graduate and Professional Student Advisory Board and the UC Committee on Affirmative Action, Diversity and Equity, among many others.

Joseph is hopeful for the anticipated impact of this award on recognizing and advancing DEI in the college.

Doreen Joseph

"As Chancellor May likes to say, I don't mind being the first, but I don't want to be the last," explained Joseph. "I'm hoping this award paves the way for the next generation of students coming into the College of Engineering and sets the stage for how the college will be moving forward."

Joseph and Esogbue met this spring at the Chancellor's Residence. In their conversation, Esogbue celebrated Joseph's selection for the award, stating multiple times, "I'm so glad we found you."

"It was a pleasure meeting Dr. Esogbue," said Joseph. "I'm incredibly honored to be part of the legacy of both Esogbue and May. They have been very encouraging, including passing the torch to myself and future award recipients to continue the important work they've both championed all these years. I don't take that responsibility lightly, and I can only hope to make as much of an impact as they both have and to carry on their legacies with excellence."  

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    University of Southern California
   
  Jun 25, 2024  
USC Catalogue 2024-2025    
USC Catalogue 2024-2025

Return to: Keck School of Medicine of USC    

Keith Administration Building 409 1975 Zonal Avenue Los Angeles, CA 90089-9031 (323) 442-1609 FAX: (323) 442-1199 Email: [email protected] Program Director: Mark R. Frey, PhD

The USC PhD Programs in Biomedical and Biological Sciences (PIBBS) is a gateway program into graduate studies at the USC Health Sciences Campus, leading to a PhD degree in a broad range of biological and biomedical disciplines.

PIBBS students experience a common first-year curriculum that will build a solid foundation in biomedical and biological sciences. Students are required to complete 29 units of core classes and laboratory rotations during the fall, spring and summer semesters of the PIBBS year, including courses in cell biology, molecular biology, human molecular genetics, protein chemistry, principles of biostatistics, ethics and accountability in biomedical research, and writing in the biomedical and biological sciences.

At the end of the spring semester of the first year, students will select a faculty adviser and a specific PhD program from among the four participating programs listed. In the second year, students will take classes that will differ depending on the PhD program they join; second year classes may include, but are not limited to, topics such as biochemistry, epidemiology, stem cell biology, cancer biology, bioinformatics, virology, immunology and physiology. In addition, each student will complete qualifying examinations for the chosen PhD program and will develop and complete an original research project that will serve as the basis for a doctoral dissertation.

PhD Graduate Programs

Cancer Biology and Genomics (PhD)     Development, Stem Cells, and Regenerative Medicine (PhD)     Infectious Diseases, Immunology and Pathogenesis (PhD)     Medical Biophysics (PhD)   Molecular Medicine (PhD)   Neuromedicine (PhD)    

PIBBS Admission Requirements

Application Deadline (priority review): December 1

Applicants to PIBBS should have a baccalaureate degree in natural sciences, or sufficient courses in mathematics and the life sciences. This is required to provide a strong background for studies in biomedical and biological research. Appropriate undergraduate degrees include biology, physiology, engineering, chemistry or computer science.

Applicants should have a strong record of academic achievement. Other requirements for admission include: a detailed statement of purpose as well as three letters of recommendation, one of which should be from a wet laboratory or computational research mentor. Previous research experience is expected. Students are admitted for the academic year in the fall semester. Applicants who are accepted with minor deficiencies are expected to correct these during the first year following enrollment. Although there is no formal application deadline, complete applications received by December 1 will be given priority.

Financial Support

Admitted students are supported by research assistantships or fellowships during their graduate career. Tuition, health insurance and health fees are also covered.

Lab Rotations

During the first year, students register for INTD 790 Research    (4 units in the fall semester and 3 units in the spring semester) and rotate through the labs of three faculty members of the program (potential research advisers). By the first summer of graduate study, but no later than after 12 months in the program, each student is expected to have selected a faculty mentor/research adviser.

PIBBS Required Core Curriculum and Research

First semester.

  • INTD 531 Cell Biology Units: 4
  • INTD 561 Molecular Biology Units: 4
  • INTD 790 Research Units: 1, 2, 3, 4, 5, 6, 7, 8, 9, 10, 11, 12

Second Semester

  • INTD 549 Protein Chemistry – Structure and Function Units: 4
  • INTD 577 Writing in the Biomedical and Biological Sciences Units: 1

Third Semester

  • INTD 500 Ethics and Accountability in Biomedical Research Units: 1
  • PM 510L Principles of Biostatistics Units: 4

Admission Requirements for PhD Programs

Admission to the Keck School of Medicine PhD program is open to all incoming PIBBS students provided all PIBBS admission requirements are met and all first year course and lab rotation requirements have been satisfactorily completed. Students from other sources, such as MD/PhD students and clinician scientists, may also be eligible on a case-by-case basis.

In general, new graduate students apply for admission to USC through the PhD programs in Biomedical and Biological Sciences (PIBBS), and become enrolled in one of four PhD programs at the Keck School of Medicine after the successful completion of the PIBBS year. During the PIBBS year, students must complete the core curriculum of 29 units, maintain a 3.0 grade point average with no grade lower than a C on all courses, complete three laboratory rotations and successfully identify a lab to join in order to continue into a PhD program.

Application information is available by contacting the PIBBS Program at [email protected] .

Unit/Course Requirements

A minimum of 60 units of graduate course credits is required for the PhD, including course work, seminars, research and dissertation units. No more than 8 units of 794 may be applied toward the PhD degree. Students must complete the first year PIBBS core curriculum as well as course requirements for their specific PhD program. Thirty units of course work, including the PIBBS core curriculum, must be completed before students are considered for the qualifying examination. Additional course work relevant to the student’s research interests  may be required by the student’s qualifying exam committee or by the student’s faculty adviser.

Guidance Committee

After 30 units of course work, which includes the PIBBS core curriculum and course requirements for one of the four PhD programs, the student, in consultation with a faculty adviser, will nominate five faculty members to serve on the guidance committee for the qualifying examination. A minimum of three of the faculty must be from the student’s PhD program, and one must be a faculty member from outside the PhD program. The chair of the guidance committee must be a member of the student’s PhD program and the faculty adviser is not allowed to be on the committee (but may be a silent presence during the exam). These nominations are submitted to the chair of the program for formal appointment.

Qualifying Examination

Students in the PhD program must pass both the written screening and the oral portions of the qualifying examination administered by their guidance committee during the second year of graduate study.

The written screening exam involves writing a research grant proposal. The deadline for completion of the written screening is the second week of January of the second year. Students who receive a failing score will be allowed one resubmission, with a deadline of within 30 days of notification of score receipt. The written portion must be passed before the oral portion can be taken.

The oral examination must be completed within 60 days after successful completion of the written screening exam. The oral examination consists of two parts. The first part consists of a presentation of the proposed thesis research. The second part consists of an open forum in which the guidance committee asks general questions on any topic related to the student’s research.

Final evaluation of the examination is determined by a consensus of the guidance committee. If a student does not pass, it is at the discretion of the committee to allow the student to repeat the oral examination within 60 days. If the student is again unable to pass, this will be grounds for dismissal from the program.

Advancement to Candidacy

Recommendation for advancement to candidacy for the PhD degree is made on the basis of the successful completion of the qualifying examination, course requirements and the student’s maintenance of at least a 3.0 GPA. A student who has not been recommended for advancement to candidacy at the end of the first semester of the third year will be dismissed from the program.

Dissertation Committee

After advancement to candidacy, the student must form a dissertation committee, in consultation with a faculty adviser. A minimum of three committee members must be selected, one of which is the faculty adviser, and at least one of which must be a tenured or tenure-track faculty member of the student’s PhD program. Committee members may be non-tenure track. The chair of the dissertation committee must be a member of the student’s PhD program and may not be the faculty adviser. The dissertation committee is responsible for counseling the student during preparation of the dissertation and conducting the final oral examination during the dissertation defense. Students are expected to meet with the dissertation committee once per year to discuss progress.

Annual Research Appraisal (ARA)

After advancing to candidacy, each graduate student presents an annual progress report to his or her dissertation research committee. Prior to the meeting, the student prepares a short written document describing significant experiments, problems and projected studies. This document is distributed to the committee and is included in the student’s file. The ARA meeting is intended to be a working session between the student and his or her committee; experimental results and problems are discussed with this context. In addition the student presents a research plan for the next year of work. A satisfactory ARA is required of every student for each year of residence after the completion of the qualifying exam. A final ARA is required the semester before the student is permitted to defend the dissertation.

Dissertation and Oral Defense

The student’s research is reported in a dissertation written under the guidance of the dissertation committee. The dissertation must demonstrate the student’s capacity for independent research, scholarly achievement and technical mastery of a special field. Students are expected to have at least one first author publication accepted in a peer-reviewed journal before the defense.

When the final draft of the dissertation is ready, the student will take the final oral defense. Students must submit their dissertations to the dissertation committee at least one month before they expect to make final revisions.

Dissertation committee members are expected to read and comment on a dissertation within two weeks from its submission. The student and faculty will coordinate a timeline for the student to present the thesis to the dissertation committee. This timeline must allow all dissertation committee members enough time to fulfill their responsibilities within the four-week deadline.

The dissertation defense is a formal public presentation of the student’s research before the program faculty and students. Dissertation defenses must be publicized at least two weeks prior to the oral defense.

All doctoral candidates must be registered in 794 Doctoral Dissertation each semester (excluding summer sessions) from the time of their advancement to candidacy until their dissertation is approved and submitted to the Graduate School.

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